English

This study examines the foraging strategies of animal resource exploitation during the Middle Preclassic period (900-300 BC) at the ancient Maya site of Pacbitun, Belize. The faunal remains analyzed in this study were recovered from various domestic structures associated with the production of shell artifacts. Detailed taphonomic analyses have revealed that the Pacbitun faunal remains were particularly affected by weathering and density-mediated attrition. White-tailed deer was the prey most frequently acquired by the Middle Preclassic Maya of Pacbitun, followed by other lower-ranked artiodactyls. A variety of less profitable prey were sometimes included in the diet breadth. Using the central place forager prey choice model as a framework, the analysis of diet breadth, habitat use, and carcass transport patterns suggests that most animal resources were acquired from terrestrial habitats, at short distances from the site. Complete carcasses of large game appear to have been frequently transported to the site, where they were exploited for their meat and marrow. Comparisons with other Middle Preclassic faunal assemblages indicate significant differences in terms of taxonomic composition, with an emphasis on the procurement of fish and turtles. It is suggested that the Middle Preclassic Maya adopted foraging strategies focusing on the exploitation of local habitats, with occasional use of exotic resources.

Author Keywords: Animal, Belize, Foraging, Maya, Subsistence, Zooarchaeology

A wind tunnel study was conducted to investigate aeolian impact ripples in sand beds of varied texture from coarsely skewed to bimodal. Experimental data is lacking for aeolian megaripples, particularly in considering the influence of wind speed on ripple morphometrics. Additionally, the modelling community requires experimental data for model validation and calibration.

Eighteen combinations of wind speed and proportion of coarse mode particles by mass were analysed for both morphometrics and optical indices of spatial segregation. Wind tunnel conditions emulated those found at aeolian megaripple field sites, specifically a unimodal wind regime and particle transport mode segregation. Remote sensing style image classification was applied to investigate the spatial segregation of the two differently coloured size populations.

Ripple morphometrics show strong dependency on wind speed. Conversely, morphometric indices are inversely correlated to the proportion of the distribution that was comprised of coarse mode particles. Spatial segregation is highly correlated to wind speed in a positive manner and negatively correlated to the proportion of the distribution that was comprised of coarse mode particles. Results reveal that the degree of spatial segregation within an impact ripple bedform can be higher than previously reported in the literature.

Author Keywords: Aeolian, Impact Ripples, Megaripple, Self-organization, Wind Tunnel

Population density regulation is a fundamental principle in ecology, however there remain several unknowns regarding the functional expression of density dependence. One prominent view is that the patterns by which density dependence is expressed are largely fixed across a species, irrespective of environmental conditions. Our study investigated the expression of density dependence in Chlamydomonas reinhartti grown under a gradient of nutrient densities, and hypothesized that the relationship between per capita growth rate (pgr) and population density would vary from concave-up to concave-down as nutrients became less limiting. Contrary to prediction, we found that the relationship between a population's pgr and density became increasingly concave-up as nutrient levels increased. Our results suggest that density dependence is strongly variable depending on exogenous and endogenous processes acting on the population, implying that expression of density regulation depends extensively on local conditions. Population growth suppression may be attributable to environments with high intraspecific competition. Additional work should reveal the mechanisms influencing how the expression of density dependence varies across populations through space and time.

Author Keywords: Chlamydomonas reinhartti, density dependence, logistic model, population dynamics, single species growth, theta-logistic equation

Inducible defenses are plastic responses by an organism to the perception of predation risk. This dissertation focuses on three experiments designed to test the hypothesis that plastic ability is limited by energetic constraints. Chapter 1 provides a general introduction to phenotypic plasticity research and the theoretical costs and limitations affecting the expression of plastic traits. In Chapter 2, I tested the hypothesis that costs of early plasticity may be manifested by a reduced response to risk in later life stages. I found that amphibian embryos are able to detect and respond to larval predators, but that the energetic cost of those plastic responses are not equivalent among behavioural, growth, and morphological traits, and their expression differs between closely-related species. Chapter 3 explicitly examines the relationship between food resource availability and plasticity in response to perceived predation risk during larval development. Food-restricted tadpoles showed limited responses to predation risk; larvae at food saturation altered behaviour, development, and growth in response to predation risk. Responses to risk varied through time, suggesting ontogeny may affect the deployment of particular defensive traits. Chapter 4 examines the influence of maternal investment into propagule size on the magnitude of the plastic responses to predation risk in resulting offspring. I found that females in better body condition laid larger eggs and that these eggs, in turn, hatched into larvae that showed greater morphological plasticity in response to predation risk. Maternal investment can therefore affect the ability of offspring to mount morphological defenses to predation risk. Last, Chapter 5 provides a synthesis of my research findings, identifying specific factors constraining the plastic responses of prey to perceived predation risk. Overall, I found constraints on plastic responses imposed by the current environment experienced by the organism (resource availability), the prior experience of the organism (predator cues in the embryonic environment), and even the condition of the previous generation (maternal body condition and reproductive investment). Together, these findings both provide new knowledge and create novel research questions regarding constraints limiting phenotypic variation in natural populations.

Author Keywords: behaviour, inducible defense, Lithobates pipiens, morphometrics, phenotypic plasticity, predation risk

Monomethylmercury (MMHg), the most bioavailable form of mercury (Hg) and a potent neurotoxin, is present at elevated concentrations in Arctic marine mammals posing serious health threats to the local populations relying on marine food for their subsistence living. The sources of MMHg in the Arctic Ocean surface water and the role of dimethylmercury (DMHg) as a source of MMHg remain unclear. The objective of this research was to determine the sources and fate of methylated Hg species (MMHg and DMHg) in the marine ecosystem by investigating processes controlling the presence of methylated Hg species in the Arctic Ocean marine boundary layer (MBL) and surface waters. A method based on solid phase adsorption on Bond Elut ENV was developed and successfully used for unprecedented measurement of methylated Hg species in the MBL in Hudson Bay (HB) and the Canadian Arctic Archipelago (CAA). MMHg and DMHg concentrations averaged 2.9 ± 3.6 (mean ± SD) and 3.8 ± 3.1 pg m-3, respectively, and varied significantly among sampling sites. MMHg in the MBL is suspected to be the product of marine DMHg degradation in the atmosphere. MMHg summer (June to September) atmospheric wet deposition rates were estimated to be 188 ± 117.5 ng m-2 and 37 ± 21.7 ng m-2 for HB and CAA, respectively, sustaining MMHg concentrations available for bio-magnification in the pelagic food web. The production and loss of methylated Hg species in surface waters was assessed using enriched stable isotope tracers. MMHg production in surface water was observed from methylation of inorganic Hg (Hg(II)) and, for the first time, from DMHg demethylation with experimentally derived rate constants of 0.92 ± 0.82 x 10-3 d-1 and 0.04 ± 0.02 d-1 respectively. DMHg demethyation rate constant (0.98 ± 0.51 d-1) was higher than that of MMHg (0.35 ± 0.25 d-1). Furthermore, relationships with environmental parameters suggest that methylated Hg species transformations in surface water are mainly biologically driven. We propose that in addition to Hg(II) methylation, the main processes controlling MMHg production in the Arctic Ocean surface waters are DMHg demethylation and deposition of atmospheric MMHg. These results are valuable for a better understanding of the cycle of methylated Hg in the Arctic marine environment.

Author Keywords: Arctic Ocean, Atmosphere, Demethylation, Dimethylmercury, Methylation, Monomethylmercury

This thesis was designed to quantify absorbance and fluorescence characteristics of dissolved organic matter (DOM) in North Atlantic, Pacific and Arctic Oceans. DOM was described in water masses of distinct sources and formation pathways as well as in regions where environmental forcings such as deep water upwelling, enhanced biological activity and receipt of freshwater discharge were prevalent. Influence of sea ice on DOM in Beaufort Sea mixed layer (0 to 30 m) seawater was investigated based on sea ice extent as well as freshwater fractions of meteoric (f_mw) and sea ice melt water (f_sim) calculated from oxygen isotope ratio (δ18O). The effect of DOM exposure to simulated solar radiation was also assessed to determine the resilience of fluorescent fractions of DOM to photodegradation. This research aims to further our ability to trace DOM in marine environments and better understand its transformation pathways and predict its fate as part of the oceanic carbon cycle in a changing climate.

Author Keywords: Absorbance, Arctic Ocean, Dissolved organic matter, Fluorescence, Parallel Factor Analysis, Sea Ice

Introduced as part of the Canada-Ontario Immigration Agreement, Local Immigration Partnerships (LIPs) mark a fundamental shift in local settlement policy. To address the gap in knowledge about the implications of this policy change, this thesis research features a case study of Durham Region's LIP. Objectives were designed to examine the impact of Durham's LIP by interviewing 52 key-informants within six sectors involved in settlement and integration. Findings indicate an effective application of the LIP policy with participants pointing to the LIP's vital role in bringing Welcome Centres to Durham, increasing the attention and profile of immigration issues and improving governance relations amongst different sectors in settlement and integration. A product of local circumstances, the LIP has engaged in a quasi-advocacy role educating mainstream service providers and institutions on how to respond to a diversifying population. Results contribute to the relatively under-studied but growing knowledge of the LIP policy while demonstrating that the localization of immigration policy under the appropriate terms can be successful.

Author Keywords: Governance, Integration, Local Immigration Partnerships, Ontario, Regionalization, Settlement

This thesis reports on the 2010 excavations of the North Group and Eastern Court at the ancient Lowland Maya site of Pacbitun. It provides a construction history of the architecture and an analysis of associated artifacts, burials, and caches. The archaeological investigations demonstrate that the seven structures (Strs. 34-40) of this restricted access plazuela group were built in the Early Classic period, and renewed in the Late Classic period. Based on analyses of artifacts (ceramics and lithics), skeletal and faunal remains, and intra- and inter-site comparisons, the North Group functioned as a secondary elite domestic residential group. Reconstruction suggests that the inhabitants here were not commoners; instead, the occupants probably were related to the ruling elite of Pacbitun. Some of the evidence includes the central location and elevation of the North Group, the presence of red painted plaster surfacing, a burial with multiple ceramic musical instruments, and multiple dedicatory caches with exotic goods (e.g., marine shell, jadeite, "Charlie Chaplin" figures).

Author Keywords: Ancient Maya, ceramic musical instruments, Charlie Chaplin figurines, Pacbitun, North Group, Eastern Court, Belize River Valley, Belize, restrictive access plazuela group, secondary elite

This thesis examines the mobilization of labour required for fortification construction during the Late Helladic (LH) period of the Aegean Bronze Age. It adopts an "energetics" approach to architecture, as a framework for systematically calculating the labour costs of construction, and using such costs to infer relative differences in political power among groups and communities through the implied differences in labour control. Accordingly, construction costs were generated for thirty-six LH fortifications, located across seven distinct regional zones of the Greek mainland and Aegean Sea. These values were then compared and evaluated against what is known of the political geographies for each region, to measure the extent to which the mobilization of labour was a function of regional power in Late Bronze Age Greece. These assessments revealed that a wide range of variation existed among the sampled regions in terms of the strength and nature of this connection, underscoring the diversity in labour relations that developed throughout the Aegean during the LH period. The labour costs were also used to suggest specific systems of recruitment that may have been in place for mobilizing workers, and to argue that fortification construction would not have been particularly burdensome or demanding for certain local populations.

Author Keywords: Energetics, Fortifications, Late Bronze Age, Monumental Architecture

The objective of this thesis is to better understand the demography and habitat selection of Newfoundland caribou. Chapter 1 provides a general introduction of elements of population ecology and behavioural ecology discussed in the thesis. In Chapter 2, I examine the causes of long-term fluctuations among caribou herds. My findings indicate that winter severity and density-dependent degradation of summer range quality offer partial explanations for the observed patterns of population change. In Chapter 3, I investigate the influence of climate, predation and density-dependence on cause-specific neonate survival. I found that when caribou populations are in a period of increase, predation from coyotes and bears is most strongly influenced by the abiotic conditions that precede calving. However, when populations begin to decline, weather conditions during calving also influenced survival. I build on this analysis in Chapter 4 by determining the influence of climate change on the interplay between predation risk and neonate survival. I found that the relative equilibrium between bears and coyotes may not persist in the future as risk from coyotes could increase due to climate change. In Chapter 5, I investigate the relationships in niche overlap between caribou and their predators and how this may influence differential predation risk by affecting encounter rates. For coyotes, seasonal changes in niche overlap mirrored variation in caribou calf risk, but had less association with the rate of encounter with calves. In contrast, changes in niche overlap during the calving season for black bears had little association with these parameters. In Chapter 6, I examine broad-level habitat selection of caribou to study trade-offs between predator avoidance and foraging during the calving season. The results suggest that caribou movements are oriented towards increased access to foraging and the reduction of encounter risk with bears, and to a lesser extent, coyotes. Finally, I synthesize the major findings from this thesis and their relevance to caribou conservation in Chapter 7, to infer that Newfoundland caribou decline is ultimately driven by extrinsic and intrinsic elements related to density-dependence. Reduction in neonate survival emerged from nutritionally-stressed caribou females producing calves with lower survival.

Author Keywords: Behavioural ecology, Black bear (Ursus americanus), Coyote (Canis latrans), Population ecology, Predator-prey interactions, Woodland caribou (Rangifer tarandus)